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Cotton & Braddy (2004) Arachnomorpha phylogeny

After Figure 8 in :

Clade 1 Clade 3 Clade 4 †Alalcomenaeus †Leanchoilia Chelicerata †Eurypterida †Weinbergina Pycnogonida †Yohoia †Jianfengia †Fortiforceps Megacheira + Chelicerata Clade 5 †Aglaspidida †Cheloniellon †Emeraldella †Lemoneites †Paleomerus †Sidneyia ‘Cheliceramorpha’ Clade 2 †Nektaspida †Buenaspis †Liwia †Soomaspis & †Tariccoia †Trilobita †Eoredlichia †Olenoides †Helmetiida †Helmetia & †Kuamaia †Saperion & †Skioldia †Tegopelte Xandarellida †Cindarella †Xandarella †Sinoburius †Retifacies ‘Trilobitomorpha’ †Burgessia Arachnomorpha Crustacea †Marrellomorpha †Marrella †Mimetaster