The evolutionary ancestry of arthropods dates back to the Cambrian period. The group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians (or their constituent clades) in a superphylum Ecdysozoa. Overall, however, the basal relationships of animals are not yet well resolved. Likewise, the relationships between various arthropod groups are still actively debated. Today, arthropods contribute to the human food supply both directly as food and more importantly, indirectly as pollinators of crops. Some species are known to spread severe disease to humans, livestock, and crops. (Full article...)
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Ammophila sabulosa taking nectar on a hemp agrimony flower head
Ammophila sabulosa, the red-banded sand wasp, is a species of the subfamily Ammophilinae of the solitary hunting wasp family Sphecidae, also called digger wasps. Found across Eurasia, the parasitoid wasp is notable for the mass provisioning behaviour of the females, hunting caterpillars mainly on sunny days, paralysing them with a sting, and burying them in a burrow with a single egg. The species is also remarkable for the extent to which females parasitise their own species, either stealing prey from nests of other females to provision their own nests, or in brood parasitism, removing the other female's egg and laying one of her own instead. (Full article...)
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The "Spanish fly", Lytta vesicatoria, has been considered to have medicinal, aphrodisiac, and other properties. Human interactions with insects include both a wide variety of uses, whether practical such as for food, textiles (especially silk), and dyestuffs, or symbolic, as in art, music, and literature, and negative interactions including damage to crops and extensive efforts to control insect pests.
Academically, the interaction of insects and society has been treated in part as cultural entomology, dealing mostly with "advanced" societies, and in part as ethnoentomology, dealing mostly with "primitive" societies, though the distinction is weak and not based on theory. Both academic disciplines explore the parallels, connections and influence of insects on human populations, and vice versa. They are rooted in anthropology and natural history, as well as entomology, the study of insects. Other cultural uses of insects, such as biomimicry, do not necessarily lie within these academic disciplines.
More generally, people make a wide range of uses of insects, both practical and symbolic. On the other hand, attitudes to insects are often negative, and extensive efforts are made to kill them. The widespread use of insecticides has failed to exterminate any insect pest, but has caused resistance to commonly used chemicals in a thousand insect species. (Full article...)
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The emerald ash borer (Agrilus planipennis), also known by the abbreviation EAB, is a green buprestid or jewel beetle native to north-eastern Asia that feeds on ash species (Fraxinus spp.). Females lay eggs in bark crevices on ash trees, and larvae feed underneath the bark of ash trees to emerge as adults in one to two years. In its native range, it is typically found at low densities and does not cause significant damage to trees native to the area. Outside its native range, it is an invasive species and is highly destructive to ash trees native to Europe and North America. Before it was found in North America, very little was known about the emerald ash borer in its native range; this has resulted in much of the research on its biology being focused in North America. Local governments in North America are attempting to control it by monitoring its spread, diversifying tree species, and through the use of insecticides and biological control. (Full article...)
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Views from above and below of male specimens from Tanzania
Pachyballus ornatus is a species of jumping spider in the genusPachyballus that lives in the forested mountains of Democratic Republic of the Congo and Tanzania. A small spider, it has a cephalothorax that is from 1.2 to 1.7 mm long and an abdomen that is between 1.7 and 2.3 mm long. The spider is generally brown with a distinctive abdominal pattern of brown patches on a yellow background. This pattern is recalled in the specific name, which means 'decorated'. Younger females are darker and lack the pattern, which appears as they mature. In an immature state, they can be confused with the related Pachyballus flavipes. The spider has yellow spinnerets and brown pedipalps. Its front legs are stouter and darker than the rest. Its copulatory organs are typical for the genus. The male has a thin coiled embolus that projects from the palpal bulb. The female has a semi-circular depression towards the front of its epigyne that is divided by a wide ridge. It was first described in 2020. (Full article...)
The samsum ant (Brachyponera sennaarensis, formerly known as Pachycondyla sennaarensis) is a species of ponerineant widely distributed in Africa and the Middle East. A common household pest, the ant is a member of the genus Brachyponera. It was first described by Austrian entomologist Gustav Mayr in 1862. The ant is known for its powerful sting, in rare occasions leading to anaphylactic shock and death. (Full article...)
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The coconut crab (Birgus latro) is a terrestrialspecies of giant hermit crab, and is also known as the robber crab or palm thief. It is the largest terrestrial arthropod known, with a weight up to 4.1 kg (9 lb). The distance from the tip of one leg to the tip of another can be as wide as 1 m (3 ft 3 in). It is found on islands across the Indian and Pacific Oceans, as far east as the Gambier Islands, Pitcairn Islands, and Caroline Island, and as far west as Zanzibar. While its range broadly shadows the distribution of the coconut palm, the coconut crab has been extirpated from most areas with a significant human population such as mainland Australia and Madagascar.
The coconut crab is the only species of the genusBirgus, and is related to the other terrestrial hermit crabs of the genus Coenobita. It shows a number of adaptations to life on land. Juvenile coconut crabs use empty gastropod shells for protection like other hermit crabs, but the adults develop a tough exoskeleton on their abdomens and stop carrying a shell. Coconut crabs have organs known as branchiostegal lungs, which they use for breathing instead of their vestigial gills. After the juvenile stage, they will drown if immersed in water for too long. They have an acute sense of smell, which they use to find potential food sources, and which has developed convergently with that of insects.
Adult coconut crabs feed primarily on fleshy fruits, nuts, seeds, and the pith of fallen trees, but they eat carrion and other organic matter opportunistically. Anything left unattended on the ground is a potential source of food, which they will investigate and may carry away – thereby getting the alternative name of "robber crab". Despite its name, coconuts are not a significant part of the crab's diet. Although it lives in a burrow, the crab has been filmed climbing coconut and pandanus trees. The crab has never been filmed selectively picking coconut fruit, though they might dislodge ripe fruit that otherwise would fall naturally. When a crab is not near its burrow, climbing is an immediate escape route from predators. Sea birds eat young crabs, and both humans and larger, older crabs eat crabs of all ages. (Full article...)
Scale insects are small insects of the orderHemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient grouping than the superfamily Coccoidea due to taxonomic uncertainties. Adult females typically have soft bodies and no limbs, and are concealed underneath domed scales, extruding quantities of wax for protection. Some species are hermaphroditic, with a combined ovotestis instead of separate ovaries and testes. Males, in the species where they occur, have legs and sometimes wings, and resemble small flies. Scale insects are herbivores, piercing plant tissues with their mouthparts and remaining in one place, feeding on sap. The excess fluid they imbibe is secreted as honeydew on which sooty mold tends to grow. The insects often have a mutualistic relationship with ants, which feed on the honeydew and protect them from predators. There are about 8,000 described species.
The oldest fossils of the group date to the Late Jurassic, preserved in amber. They were already substantially diversified by the Early Cretaceous suggesting an earlier origin during the Triassic or Jurassic. Their closest relatives are the jumping plant lice, whiteflies, phylloxera bugs and aphids. The majority of female scale insects remain in one place as adults, with newly hatched nymphs, known as "crawlers", being the only mobile life stage, apart from the short-lived males. The reproductive strategies of many species include at least some amount of asexual reproduction by parthenogenesis.
Some scale insects are serious commercial pests, notably the cottony cushion scale (Icerya purchasi) on Citrus fruit trees; they are difficult to control as the scale and waxy covering protect them effectively from contact insecticides. Some species are used for biological control of pest plants such as the prickly pear, Opuntia. Others produce commercially valuable substances including carmine and kermes dyes, and shellac lacquer. The two red colour-names crimson and scarlet both derive from the names of Kermes products in other languages. (Full article...)
Chamberlin was a prolific taxonomist who named over 4,000 new animal species in over 400 scientific publications. He specialized in arachnids (spiders, scorpions, and relatives) and myriapods (centipedes, millipedes, and relatives), ranking among the most prolific arachnologists and myriapodologists in history. He described over 1,400 species of spiders, 1,000 species of millipedes, and the majority of North American centipedes, although the quantity of his output was not always matched with quality, leaving a mixed legacy to his successors. He also did pioneering ethnobiological studies with the Goshute and other indigenous people of the Great Basin, cataloging indigenous names and cultural uses of plants and animals. Chamberlin was celebrated by his colleagues at the University of Utah, however he was disliked among some arachnologists, including some of his former students. After retirement he continued to write, publishing on the history of education in his home state, especially that of the University of Utah.
Chamberlin was a member of the Church of Jesus Christ of Latter-day Saints (LDS Church). In the early twentieth century, Chamberlin was among a quartet of popular Mormon professors at Brigham Young University whose teaching of evolution and biblical criticism resulted in a 1911 controversy among University and Church officials, eventually resulting in the resignation of him and two other professors despite widespread support from the student body, an event described as Mormonism's "first brush with modernism". (Full article...)
Among parasitoids, strategies range from living inside the host (endoparasitism), allowing it to continue growing before emerging as an adult, to paralysing the host and living outside it (ectoparasitism). Hosts can include other parasitoids, resulting in hyperparasitism; in the case of oak galls, up to five levels of parasitism are possible. Some parasitoids influence their host's behaviour in ways that favour the propagation of the parasitoid.
Male Unicorn sikus, viewed from above (legs omitted).
Unicorn ("one horn", in Latin) is a genus of goblin spiders (family Oonopidae) from South America, containing seven species that occur predominantly in high elevation, semi-desert regions of Bolivia, Chile, and Argentina. Individuals are relatively large for goblin spiders, measuring up to 3.0 mm (0.12 in) in body length. The genus name refers to a characteristic pointed projection between the eyes and jaws of males. In at least one species, broken-off tips of the male pedipalps have been found within the genitalia of females, postulated as a means of sperm competition. Unicorn possesses several traits that suggest it is a relatively "primitive" member of the Oonopidae, and is classified with other similar, soft-bodied goblin spiders in the subfamily Sulsulinae. (Full article...)
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Adult male
The housefly (Musca domestica) is a fly of the suborder Cyclorrhapha. It possibly originated in the Middle East, and spread around the world as a commensal of humans. Adults are gray to black, with four dark, longitudinal lines on the thorax, slightly hairy bodies, and a single pair of membranous wings. They have red compound eyes, set farther apart in the slightly larger female.
The female housefly usually mates only once and stores the sperm for later use. It lays batches of about 100 eggs on decaying organic matter such as food waste, carrion, or feces. These soon hatch into legless white larvae, known as maggots. After two to five days of development, these metamorphose into reddish-brown pupae, about 8 millimetres (3⁄8 inch) long. Adult flies normally live for two to four weeks, but can hibernate during the winter. The adults feed on a variety of liquid or semi-liquid substances, as well as solid materials which have been softened by their saliva. They can carry pathogens on their bodies and in their feces, contaminate food, and contribute to the transfer of food-borne illnesses, while, in numbers, they can be physically annoying. For these reasons, they are considered pests.
Houseflies, with short life cycles and ease with which they can be maintained, have been found useful for laboratory research into aging and sex determination. Houseflies appear in literature from Ancient Greek myth and Aesop's "The Impertinent Insect" onwards. Authors sometimes choose the housefly to speak of the brevity of life, as in William Blake's 1794 poem "The Fly", which deals with mortality subject to uncontrollable circumstances. (Full article...)
Earwigs make up the insectorderDermaptera. With about 2,000 species in 12 families, they are one of the smaller insect orders. Earwigs have characteristic cerci, a pair of forceps-like pincers on their abdomen, and membranous wings folded underneath short, rarely used forewings, hence the scientific order name, "skin wings". Some groups are tiny parasites on mammals and lack the typical pincers. Earwigs are found on all continents except Antarctica.
Earwigs are mostly nocturnal and often hide in small, moist crevices during the day, and are active at night, feeding on a wide variety of insects and plants. Damage to foliage, flowers, and various crops is commonly blamed on earwigs, especially the common earwig Forficula auricularia.
Earwigs have five molts in the year before they become adults. Many earwig species display maternal care, which is uncommon among insects. Female earwigs may care for their eggs; the ones that do will continue to watch over nymphs until their second molt. As the nymphs molt, sexual dimorphism such as differences in pincer shapes begins to show. (Full article...)
Metanephrops challengeri (commonly known as the New Zealand lobster or New Zealand scampi) is a species of slim, pink lobster that lives around the coast of New Zealand. It is typically 13–18 cm (5–7 in) long and weighs around 100 g (3.5 oz). The carapace and abdomen are smooth, and adults are white with pink and brown markings and a conspicuous pair of long, slim claws. M. challengeri lives in burrows at depths of 140–640 m (460–2,100 ft) in a variety of sediments. Although individuals can live for up to 15 years, the species shows low fecundity, where small numbers of larvae hatch at an advanced stage.
M. challengeri is a significant prey item for ling, as well as being an important fishery species for human consumption; trawlers catch around 1,000 t (2,200,000 lb) per year under the limitations of New Zealand's Quota Management System. The species was first collected by the Challenger expedition of 1872–1876, but only described as separate from related species by Heinrich Balss in 1914. Although originally classified in the genus Nephrops, it was moved in 1972 to a new genus, Metanephrops, along with most other species then classified in Nephrops. (Full article...)
Myrmecia is a genus of ants first established by Danish zoologistJohan Christian Fabricius in 1804. The genus is a member of the subfamily Myrmeciinae of the family Formicidae. Myrmecia is a large genus of ants, comprising at least 93 species that are found throughout Australia and its coastal islands, while a single species is only known from New Caledonia. One species has been introduced out of its natural distribution and was found in New Zealand in 1940, but the ant was last seen in 1981. These ants are commonly known as bull ants, bulldog ants or jack jumper ants, and are also associated with many other common names. They are characterized by their extreme aggressiveness, ferocity, and painful stings. Some species are known for the jumping behavior they exhibit when agitated.
Species of this genus are also characterized by their elongated mandibles and large compound eyes that provide excellent vision. They vary in colour and size, ranging from 8 to 40 millimetres (0.31 to 1.57 in). While workers and queens are hard to distinguish from each other due to their similar appearance, males are identifiable by their perceptibly smaller mandibles. Almost all Myrmecia species are monomorphic, with little variation among workers of a given species. Some queens are ergatoid and have no wings, while others have either stubby or completely developed wings. Nests are mostly found in soil, but they can be found in rotten wood and under rocks. One species does not nest in the ground at all; its colonies can only be found in trees.
A queen will mate with one or more males, and during colony foundation she will hunt for food until the brood have fully developed. The life cycle of the ant from egg to adult takes several months. Myrmecia workers exhibit greater longevity in comparison to other ants, and workers are also able to reproduce with male ants. Myrmecia is one of the most primitive group of ants on earth, exhibiting differentiated behaviors from other ants. Workers are solitary hunters and do not lead other workers to food. Adults are omnivores that feed on sweet substances, but the larvae are carnivores that feed on captured prey. Very few predators eat these ants due to their sting, but their larvae are often consumed by blindsnakes and echidnas, and a number of parasites infect both adults and brood. Some species are also effective pollinators. (Full article...)
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female
Afraflacilla braunsi is a species of jumping spider in the genusAfraflacilla. First found in South Africa, the spider was subsequently observed living in Saudi Arabia, Turkmenistan, the United Arab Emirates and Yemen, although it is likely to have a wider distribution. First described in 1903 by George and Elizabeth Peckham, it was originally allocated to the genus Pseudicius with the name Pseudicius braunsii. After being renamed Icius braunsi in 1987, it was finally given its current name by Jerzy Prószyński in 2017. Pseudicius tripunctatus, now called Afraflacilla tripunctatus, is a synonym.
A small to medium-sized spider, Afraflacilla braunsi has a carapace that is between 1.7 and 1.9 mm (0.07 and 0.07 in) long and an abdomen 2.2 and 3.2 mm (0.09 and 0.13 in) long. The carapace is brown, sometimes with yellow sides. The abdomen is light brown or fawn with lighter yellow patches. It has large brown front legs, the remainder generally yellow and less robust. It makes noises by rubbing its forelegs against small hairs under its eyes. The copulatory organs are distinctive for the species. The male has a characteristically bulbous palpal bulb and a single long projection, or apophysis, extending from the palpal tibia. The female has very long and coiled insemination ducts that lead to large spermathecae. (Full article...)
The following are images from various arthropod-related articles on Wikipedia.
Image 1Ghost crab, showing a variety of integument types in its exoskeleton, with transparent biomineralization over the eyes, strong biomineralization over the pincers, and tough chitin fabric in the joints and the bristles on the legs (from Arthropod exoskeleton)
Image 3The house centipedeScutigera coleoptrata has rigid sclerites on each body segment. Supple chitin holds the sclerites together and connects the segments flexibly. Similar chitin connects the joints in the legs. Sclerotised tubular leg segments house the leg muscles, their nerves and attachments, leaving room for the passage of blood to and from the hemocoel (from Arthropod exoskeleton)
Image 9Formation of anterior segments across arthropod taxa based on gene expression and neuroanatomical observations, Note the chelicera(Ch) and chelifore(Chf) arose from somite 1 and thus correspond to the first antenna(An/An1) of other arthropods. (from Chelicerata)
Image 10The fangs in spiders' chelicerae are so sclerotised as to be greatly hardened and darkened (from Arthropod exoskeleton)
Image 12In honeypot antrepletes, the abdomens of the workers that hold the sugar solution grow vastly, but only the unsclerotised cuticle can stretch, leaving the unstretched sclerites as dark islands on the clear abdomen (from Arthropod exoskeleton)
Image 17Mature queen of a termite colony, showing how the unsclerotised cuticle stretches between the dark sclerites that failed to stretch as the abdomen grew to accommodate her ovaries (from Arthropod exoskeleton)
Image 18Some of the various hypotheses of myriapod phylogeny. Morphological studies (trees a and b) support a sister grouping of Diplopoda and Pauropoda, while studies of DNA or amino acid similarities suggest a variety of different relationships, including the relationship of Pauropoda and Symphyla in tree c. (from Myriapoda)
Image 19Crab larva barely recognisable as a crab, radically changes its form when it undergoes ecdysis as it matures (from Arthropod exoskeleton)
Image 22 This fully-grown robber crab has tough fabric forming its joints, delicate biomineralized cuticle over its sensory antennae, optic-quality over its eyes, and strong, calcite-reinforced chitin armouring its body and legs; its pincers can break into coconuts (from Arthropod exoskeleton)
Image 24This Zoea-stage larva is hardly recognisable as a crab, but each time it sheds its cuticle it remodels itself, eventually taking on its final crab form (from Arthropod exoskeleton)
Image 25Decapods, from Ernst Haeckel's 1904 work Kunstformen der Natur (from Crustacean)
A honey bee extracts nectar from a flower using its proboscis. Tiny hairs covering the bee's body maintain a slight electrostatic charge, causing pollen from the flower's anthers to stick to the bee's hairs, allowing for pollination when the bee moves on to another flower.
The Forest scorpion (Cercophonius squama) is a scorpion native to southeastern Australia and Tasmania. The body is 25 to 40 millimetres (0.98 to 1.57 in) long, and coloured creamy yellow to orange brown with dark brown variegations. The legs are yellow with some dark brown pigment.
The Globe Skimmer (Pantala flavescens) is the most widespread dragonfly species on the planet, found between about the 40th parallels of latitude, or where the annual mean temperature is above 20 °C (68 °F), except in Europe where there are only occasional sightings.
The citrus root weevil (Diaprepes abbreviatus) is a major agricultural pest weevil in several Caribbean countries and the U.S. state of Florida. The larvae feed on the roots of the host plant for several months. They often eat the taproot of the plant, which can kill it by depriving it of water and nutrients or by making it vulnerable to infection by fungi, or water moulds.
Xylotrupes socrates (Siamese rhinoceros beetle, or "fighting beetle"), male, on a banana leaf. This scarab beetle is particularly known for its role in insect fighting in Northern Laos and Thailand.
The orb-weaver spiders (family Araneidae) are the familiar builders of spiral wheel-shaped webs often found in gardens, fields and forests. The family is a large one, including over 2800 species in over 160 genera worldwide, making it the third largest known (behind Salticidae and Linyphiidae). The web has always been thought of as an engineering marvel.
The Ozyptila praticola species of crab spider is found throughout Europe and the Middle East. They do not build webs to trap prey, but are active hunters. Crab spiders are so named because of their first two pairs of legs, which are held out to the side giving them a crab-like appearance. Also, like crabs, these spiders move sideways and backwards more easily than forwards.
A compound eye is a visual organ found in certain arthropods. The compound eye consists of between 12 and 1,000 ommatidia, little dark/bright sensors. The image perceived by the arthropod is "recalculated" from the numerous ommatidia which point in slightly different directions. In contrast to other eye types, there is no central lens or retina. Though the resulting image is poor in resolution, it can detect quick movements and, in some cases, the polarization of light.
Dragonflies have about 30,000 facets to their compound eyes, giving them nearly a 360° field of vision.
Gonepteryx rhamni, also known as the common brimstone, is a butterfly in the family Pieridae. It lives throughout the Palearctic zone and is commonly found across Europe, Asia and North Africa. The butterfly relies on two species of buckthorn as hosts for its eggs and larvae, which influences its geographic range and distribution as these plants are commonly found in wetlands. After spending the summer feeding, adults travel to woodland areas to spend seven months hibernating. In spring when their host plants have developed, they return to the wetlands to breed and lay eggs. Both the larval and adult forms of the species have protective coloration and behaviour that decreases their chances of being recognised and preyed upon. The adult common brimstone has sexual dichromism in its wing coloration and iridescence; the male (pictured) has yellow wings and iridescence, while females have greenish-white wings and are not iridescent.
A macro view of a Gonia capitatafly feeding on honey, showing its proboscis and pedipalps (the two appendages protruding from the proboscis), two types of insect mouthparts. The proboscis actually comprises the labium, a quadrupedal structure, and a sponge-like labellum at the end. Flies eat solid food by secreting saliva and dabbing it over the food item. As the saliva dissolves the food, the solution is then drawn up into the mouth as a liquid. The labellum's surface is covered by minute food channels which form a tube leading to the esophagus, and food is drawn up the channels by capillary action.
The Meadow Argus (Junonia villida) is a species of butterfly native to Australasia. Its brown wings are each covered with two distinctive black and blue eyespots as well as white and orange marks that appear on the edge of the wings. Males and females are similar in appearance and size, with females being slightly larger.
A lithograph by Sidney Hall, published in Urania's Mirror, depicting the constellation Scorpius. One of the 48 constellations identified by the Greek astronomer Ptolemy in the second century, Scorpius lies between Libra to the west and Sagittarius to the east. It contains 18 main stars, the brightest of which is Antares. Although frequently identified as a scorpion, this constellation has also been likened to a "leaning coconut tree" (Java) or "Maui's fishhook" (Hawaii).
The light blue soldier crab (Mictyris longicarpus) inhabits beaches in the Indo-Pacific region. Soldier crabs filter sand or mud for microorganisms. They congregate during the low tide, and bury themselves in a corkscrew pattern during high tide, or whenever they are threatened.
The oriental hornet (Vespa orientalis) is a social insect in the family Vespidae, found in southern Europe, northeastern Africa, and western Asia. This worker, photographed in Sha'ar Poleg Reserve in Israel, is gathering nectar from a sea squill; adults will also feed on honeydew and fruit. They also capture insects such as grasshoppers, flies and honey bees to provide a diet high in protein for the colony's brood.
Danaus genutia, the common tiger or striped tiger, is a species of brush-footed butterfly found in Sri Lanka, India, Myanmar, south-eastern Asia and Australia. It prefers areas of moderate to heavy rainfall, and typical habitats include scrubby jungle, deciduous forests and fallow land near habitations. The insect sequesters toxins from plants, and advertises its unpalatability by having prominent markings and striking colour patterns. This adult male common tiger, of the subspecies D. g. genutia, was photographed in Kerala, India.
The red rock crab (Grapsus grapsus), also known as "Sally Lightfoot", is one of the commonest crabs along the western seaboard of the Americas. John Steinbeck wrote of them, "Everyone who has seen them has been delighted with them ... These little crabs, with brilliant cloisonné carapaces, walk on their tiptoes, they have remarkable eyes and an extremely fast reaction time." He tried to catch them but to little avail. "If you walk slowly, they move slowly ahead of you in droves. If you hurry, they hurry. When you plunge at them, they seem to disappear in a puff of blue smoke."
Planthoppers are insects in the suborder Auchenorrhyncha. This photograph shows three adult Phromnia rosea planthoppers on a stem, with three nymphs underneath; the adults fold their wings in a tent-like fashion, while the nymphs are clad in a dense tangle of white wax threads. Both the adults and the nymphs feed by sucking sap from the host plant.
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